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E Graph of extension vs. The extension plotted here is the extension for zero gape subtracted from the extension for tyrannosaurid paleobiology given gape, so that the extension for zero gape is zero.

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This graph therefore represents the relation of gape to the length of the muscle extended beyond its tyrannosaurid paleobiology rest length with the mouth closed.

The units of the abscissa are centimeters times 0.


The points on this graph tyrannosaurid paleobiology B, C, and D are likewise derived from the extensions illustrated in parts BC tyrannosaurid paleobiology, and Drespectively. F Graph of the lever arm versus gape.

This graph is constructed from the lengths of the lever arms as tyrannosaurid paleobiology in BCand D. The abscissa represents the lever arm, and the ordinate the tyrannosaurid paleobiology.

Matthews Find articles by Neffra A. George Pemberton Find articles by S.

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The authors have declared that no competing interests exist. Conceived and designed the experiments: Contributed to the writing of the tyrannosaurid paleobiology Collected measured specimen data: Received Apr 9; Accepted Jul 2.


Copyright notice This is an open-access article distributed under the terms of the Creative Commons Public Domain declaration, which stipulates tyrannosaurid paleobiology, once placed in the public domain, this work may be freely reproduced, distributed, transmitted, modified, built upon, or otherwise tyrannosaurid paleobiology by anyone for any lawful purpose.

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This article has been cited by other articles in PMC. Associated Data Data Availability Statement The authors confirm that all data underlying the findings are fully available without restriction. All data on the newly described tyrannosaurid trackways are available in Table 1 of this manuscript.

All tyrannosaurid paleobiology used in the calculations of size, speed, and age were previously used in Currie PJ Tyrannosaurid paleobiology growth in tyrannosaurids Dinosauria: Can J Earth Sci Abstract The skeletal record of tyrannosaurids is well-documented, whereas their footprint record is surprisingly sparse.